Cloeon dipterum

COMPLETE = 4.0; MRE = 0.161; SMAE = 0.184; SMSE = 0.049

Data & predictions

data prd RE symbol units description
10 8.109 0.1891 ab d age at birth
30 34.74 0.1581 tj d time at emergence
21 20.1 0.04279 am d life span as imago
23.6 26.53 0.124 tp d time since birth at puberty
6.4 8.217 0.284 tpj d time at emergence since puberty
111.6 109.7 0.01646 tp_c d time since birth at puberty
144 134.8 0.06436 tj_c d time at emergence
0.0167 0.01399 0.1623 L0 cm egg length
0.0554 0.05195 0.06236 Lb cm length at hatching
0.6845 0.5505 0.1958 Lp cm body length female end of instar 6
0.92 0.7713 0.1616 Lj cm body length female sub-imago
0.82 0.7691 0.06208 Lj_m cm body length male sub-imago
1.03 0.6656 0.3538 Lj_w cm wing length female imago
0.0006 0.001306 1.176 Wd0 mg initial dry weight
0.0009 0.0004765 0.4705 Wdb mg dry weight at birth
13.6 7.976 0.4135 Wwj mg wet weight female subimago
8.2 7.902 0.0363 Wwj_m mg wet weight male subimago
0.1962 0.1801 0.08214 L1 cm body length of instar 1
0.263 0.2465 0.06292 L2 cm body length of instar 2
0.3654 0.3949 0.08074 L3 cm body length of instar 3
0.4988 0.4164 0.1651 L4 cm body length of instar 4
0.6015 0.5565 0.07488 L5 cm body length of instar 5
0.6845 0.7232 0.05648 L6 cm body length of instar 6
0.6703 0.8304 0.2389 L7 cm body length of instar 7
0.6745 0.9095 0.3484 L8 cm body length of Nymph instar 1
0.6695 0.9432 0.4088 L9 cm body length of Nymph instar 2
0.02078 0.01985 0.04454 Wd1 mg dry weight of instar 1
0.06111 0.05089 0.1672 Wd2 mg dry weight of instar 2
0.1175 0.2094 0.7812 Wd3 mg dry weight of instar 3
0.3216 0.2455 0.2367 Wd4 mg dry weight of instar 4
0.8554 0.5858 0.3152 Wd5 mg dry weight of instar 5
1.091 1.286 0.1779 Wd6 mg dry weight of instar 6
1.105 1.342 0.2147 Wd7 mg dry weight of instar 7
1.024 1.347 0.3144 Wd8 mg dry weight of Nymph instar 1
0.9199 0.6293 0.3159 Wd9 mg dry weight of Nymph instar 2
0.033 0.02374 0.2805 Lh1 cm head length of instar 1
0.0422 0.03249 0.23 Lh2 cm head length of instar 2
0.0584 0.05206 0.1085 Lh3 cm head length of instar 3
0.0732 0.0549 0.25 Lh4 cm head length of instar 4
0.0992 0.07336 0.2605 Lh5 cm head length of instar 5
0.1092 0.09534 0.1269 Lh6 cm head length of instar 6
0.1153 0.1095 0.05044 Lh7 cm head length of instar 7
0.1092 0.1199 0.09807 Lh8 cm head length of Nymph instar 1
0.1153 0.1244 0.07851 Lh9 cm head length of Nymph instar 2
4.6 3.258 0.2917 t5 d duration of instar 5
5.3 3.424 0.354 t6 d duration of instar 6
3 2.334 0.222 t7 d duration of instar 7
2.3 1.911 0.1692 t8 d duration of instar 8
1.1 0.9158 0.1675 t9 d duration of instar 9
1.683e-05 1.886e-05 0.1204 JX5 g food uptake instar 5
4.175e-05 3.904e-05 0.06487 JX6 g food uptake instar 6
2.577e-05 2.341e-05 0.09162 JX7 g food uptake instar 7
1.485e-05 1.597e-05 0.07543 JX8 g food uptake instar 8
4.5e-07 4.802e-07 0.06704 JX9 g food uptake instar 9
463.9 446.7 0.03697 N_HFT # eggs/female at High Food Treatment
319.4 356.7 0.1165 N_MFT # eggs/female at Medium Food Treatment
216.7 226.7 0.04649 N_LFT # eggs/female at Low Food Treatment
see figure 0.2773 tL_f time vs. length
see figure 0.2778 tL_m time vs. length
see figure 0.2629 tW_f time vs. wet weight
see figure 0.2605 tW_m time vs. wet weight
see figure 0.3929 LJO_f length vs. O_2 consumption
see figure 0.3915 LJO_m length vs. O_2 consumption
see figure 0.1434 LWw_f length vs. wet weight
see figure 0.1441 LWw_m length vs. wet weight
see figure 0.1353 tL_f1 time since start of experiment vs. length
see figure 0.1424 tL_m1 time since start of experiment vs. length
see figure 0.05718 tL_c time vs. length
see figure 0.06758 tL_w time vs. length
see figure 0.07468 tL_g time vs. length
see figure 0.2745 Ttj_g Temperature vs. time since birth at emergence
see figure 0.05633 tL_h1 time since start of experiment vs. head length
see figure 0.05601 tL_h2 time since start of experiment vs. head length
see figure 0.04776 tL_h3 time since start of experiment vs. head length
see figure 0.2546 tL_h4 time since start of experiment vs. head length
see figure 0.0357 tL_h5 time since start of experiment vs. head length
see figure 0.2277 Lw_Wd wing length female imago vs. dry weight
see figure 0.2893 Lw_N wing length female imago vs. No. of hatchlings
see figure 0.3669 Ttj Temperature vs. time since birth at emergence
see figure 0.09937 tS starvation time vs. survival
see figure 0.09811 tS_2 starvation time vs. survival
see figure 0.03005 tS_3 starvation time vs. survival
see figure 0.2349 WJO_T1 wet weight vs. oxygen consumption
see figure 0.1922 WJO_T2 wet weight vs. oxygen consumption
see figure 0.3092 WJO_T3 wet weight vs. oxygen consumption
see figure 0.1247 WJO_T4 wet weight vs. oxygen consumption
see figure 0.2222 WJO_T5 wet weight vs. oxygen consumption
see figure 0.1163 L_Wd body length female imago vs. dry weight
0.02 0.004306 0.7847 psd.v cm/d energy conductance
18 181.8 9.101 psd.p_M J/d.cm^3 vol-spec som maint
0.002 0.03082 14.41 psd.k_J 1/d maturity maint rate coefficient
0.99 0.9994 0.009539 psd.k - maintenance ratio
0.8 0.72 0.09995 psd.kap - allocation fraction to soma
0.8 0.6032 0.246 psd.kap_G - growth efficiency
0.95 0.95 0 psd.kap_R - reproduction efficiency

hep parameters at 20.0°C

symbol units value free description
T_ref K 293.1 0 Reference temperature
T_A K 8234 0 Arrhenius temperature
z - 0.2915 0 zoom factor for female
F_m l/d.cm^2 6.5 0 {F_m}, max spec searching rate
kap_X - 0.8 0 digestion efficiency of food to reserve
kap_P - 0.1 0 faecation efficiency of food to faeces
v cm/d 0.004306 0 energy conductance
kap - 0.72 0 allocation fraction to soma
kap_R - 0.95 0 reproduction efficiency
p_M J/d.cm^3 181.8 0 [p_M], vol-spec somatic maint
p_T J/d.cm^2 0 0 {p_T}, surf-spec somatic maint
k_J 1/d 0.03082 0 maturity maint rate coefficient
E_G J/cm^3 5896 0 [E_G], spec cost for structure
E_Hb J 0.002663 0 maturity at birth
E_Hp J 3.169 0 maturity at puberty
E_Rj J/cm^3 2032 0 reproduction buffer density at emergence
h_a 1/d^2 0.002204 0 Weibull aging acceleration
s_G - 0.0001 0 Gompertz stress coefficient
L_0 cm 0.03302 0 initial length for gaia2019 head length data
L_0_f cm 0.2146 0 initial length for gaia2016 data
L_0_m cm 0.218 0 initial length for gaia2016 data
L_ini_W cm 0.652 0 initial length tS (Wage2015)
L_ini_c cm 0.2678 0 initial length tL_c (gaia2016, winter)
L_ini_g cm 0.3574 0 initial length tL (gaia2016, summer)
L_ini_w cm 0.2678 0 initial length tL_w (gaia2016, winter)
T_AL K 7.172e+04 0 Arrhenius temperature for lower boundary
T_L K 283.9 0 lower boundary temperature
dV need to put 0.0764 0 scalar with specific density in g/cm^3 of dry mass
del_M - 0.2024 0 shape coefficient
del_Mh - 1.535 0 shape coefficient for head length
del_Mw - 0.2868 0 shape coefficient for wing length
del_Y - 1.41 0 shape coefficient for egg
e_critical - 0.2089 1 critical reserve density
f - 1 0 scaled functional response for 0-var data
f_7 - 0.598 0 scaled functional responce for instar 7
f_8 - 0.3979 0 scaled functional responce for instar 8
f_9 - 0.02205 0 scaled functional responce for instar 9
f_HFT - 1 0 scaled functional response for HFT
f_LFT - 0.1183 0 scaled functional response for LFT
f_MFT - 0.3221 0 scaled functional response for MFT
f_S - 0.4806 0 scaled functional response for Swee2018 data
f_W - 1 0 scaled functional response (Wage2015)
f_c - 0.1324 0 scaled functional response for Cian1979 data
f_fm1 - 0.8228 0 scaled functional response for tL data (gaia2016)
f_g - 1 0 scaled functional response for gaia2019 data
f_g13 - 0.5165 0 scaled functional response for gaia2019 data, 13oC
f_g18 - 0.6601 0 scaled functional response for gaia2019 data, 18oC
f_g22 - 0.6383 0 scaled functional response for gaia2019 data, 22oC
f_g26 - 0.4348 0 scaled functional response for gaia2019 data, 26oC
f_g8 - 0.5948 0 scaled functional response for gaia2019 data, 8oC
f_tL_c - 0.2392 0 scaled functional response for tL data (gaia2016 semifield winter, cold)
f_tL_g - 0.1743 0 scaled functional response for tL data (gaia2016 semifield summer)
f_tL_w - 0.1539 0 scaled functional response for tL data (gaia2016 semifield winter, warm)
h_b 1/d 0.002929 1 background hazard rate (Wage2015)
s_1 - 12.02 0 stress at instar 1: L_1^2/ L_b^2
s_2 - 1.873 0 stress at instar 2: L_2^2/ L_1^2
s_3 - 2.567 0 stress at instar 3: L_3^2/ L_2^2
s_4 - 1.112 0 stress at instar 4: L_4^2/ L_3^2
s_5 - 1.786 0 stress at instar 5: L_5^2/ L_4^2
s_6 - 1.689 0 stress at instar 6: L_6^2/ L_5^2
s_7 - 1.319 0 stress at instar 7 (N1): L_7^2/ L_6^2
s_8 - 1.2 0 stress at instar 8 (N2): L_8^2/ L_7^2
k_starv 1/d 0.2183 1 starvation rate coefficient
tau_1 - 2.209 0 scaled time related to time-variable f: end of decrease
y_cl g/mol 0.364 0 conversion from mol of periphyton to g of chlorophyl
z_m - 0.2907 0 zoom factor for male