Lumbricus rubellus allStat version: 2022/02/27
model - std NA NA typified model
T_typical °C 22.5 NA NA typical body temperature
T_ref K 293.15 NA NA reference temperature
T_A K 8000 NA NA Arrhenius temperature
z - 0.91592 NA NA zoom factor
F_m l/d.cm^2 6.5 20 NA {F_m}, max spec searching rate
kap_X - 0.8 NA NA digestion efficiency of food to reserve
kap_P - 0.1 NA NA faecation efficiency of food to faeces
v cm/d 0.023367 20 NA energy conductance
kap - 0.98502 NA NA allocation fraction to soma
kap_R - 0.475 NA NA reproduction efficiency
p_M J/d.cm^3 371.632 20 NA [p_M], vol-spec somatic maint
p_T J/d.cm^2 0 20 NA {p_T}, surf-spec somatic maint
k_J 1/d 0.002 20 NA maturity maint rate coefficient
E_G J/cm^3 4182.94 NA NA [E_G], spec cost for structure
E_Hb J 0.1412 NA NA maturity at birth
E_Hp J 38.78 NA NA maturity at puberty
h_a 1/d^2 4.751e-09 20 NA Weibull aging acceleration
s_G - 0.0001 NA NA Gompertz stress coefficient
d_X g/cm^3 0.16 NA NA specific density of food
d_V g/cm^3 0.16 NA NA specific density of structure
d_E g/cm^3 0.16 NA NA specific density of reserve
d_P g/cm^3 0.16 NA NA specific density of faeces
mu_X J/ mol 525000 NA NA chemical potential of food
mu_V J/ mol 500000 NA NA chemical potential of structure
mu_E J/ mol 550000 NA NA chemical potential of reserve
mu_P J/ mol 480000 NA NA chemical potential of faeces
mu_C J/ mol 0 NA NA chemical potential of CO2
mu_H J/ mol 0 NA NA chemical potential of H2O
mu_O J/ mol 0 NA NA chemical potential of O2
mu_N J/ mol 122000 NA NA chemical potential of N-waste
n_CX - 1 NA NA chem. index of carbon in food
n_HX - 1.8 NA NA chem. index of hydrogen in food
n_OX - 0.5 NA NA chem. index of oxygen in food
n_NX - 0.15 NA NA chem. index of nitrogen in food
n_CV - 1 NA NA chem. index of carbon in structure
n_HV - 1.8 NA NA chem. index of hydrogen in structure
n_OV - 0.5 NA NA chem. index of oxygen in structure
n_NV - 0.15 NA NA chem. index of nitrogen in structure
n_CE - 1 NA NA chem. index of carbon in reserve
n_HE - 1.8 NA NA chem. index of hydrogen in reserve
n_OE - 0.5 NA NA chem. index of oxygen in reserve
n_NE - 0.15 NA NA chem. index of nitrogen in reserve
n_CP - 1 NA NA chem. index of carbon in faeces
n_HP - 1.8 NA NA chem. index of hydrogen in faeces
n_OP - 0.5 NA NA chem. index of oxygen in faeces
n_NP - 0.15 NA NA chem. index of nitrogen in faeces
n_CC - 1 NA NA chem. index of carbon in CO2
n_HC - 0 NA NA chem. index of hydrogen in CO2
n_OC - 2 NA NA chem. index of oxygen in CO2
n_NC - 0 NA NA chem. index of nitrogen in CO2
n_CH - 0 NA NA chem. index of carbon in H2O
n_HH - 2 NA NA chem. index of hydrogen in H2O
n_OH - 1 NA NA chem. index of oxygen in H2O
n_NH - 0 NA NA chem. index of nitrogen in H2O
n_CO - 0 NA NA chem. index of carbon in O2
n_HO - 0 NA NA chem. index of hydrogen in O2
n_OO - 2 NA NA chem. index of oxygen in O2
n_NO - 0 NA NA chem. index of nitrogen in O2
n_CN - 1 NA NA chem. index of carbon in N-waste
n_HN - 2 NA NA chem. index of hydrogen in N-waste
n_ON - 1 NA NA chem. index of oxygen in N-waste
n_NN - 2 NA NA chem. index of nitrogen in N-waste
c_T - 1.25956 22.5 NA Temperature Correction factor
p_Am J/d.cm^2 345.562 20 NA {p_Am}, spec assimilation flux
w_X g/mol 23.9 NA NA molecular weight for water-free food
w_V g/mol 23.9 NA NA molecular weight for water-free structure
w_E g/mol 23.9 NA NA molecular weight for water-free reserve
w_P g/mol 23.9 NA NA molecular weight for water-free product (feaces)
M_V mol/cm^3 0.00669456 NA NA [M_V], volume-specific mass of structure
y_V_E mol/mol 0.880244 NA NA yield of structure on reserve
y_E_V mol/mol 1.13605 NA NA yield of reserve on structure
k_M 1/d 0.0888448 20 NA somatic maintenance rate coefficient
k - 0.0225112 NA NA maintenance ratio
E_m J/cm^3 14788.5 NA NA [E_m], reserve capacity
m_Em mol/mol 4.01641 NA NA reserve capacity
g - 0.287153 NA NA energy investment ratio
L_m cm 0.91592 NA NA maximum structural length
L_T cm 0 NA NA heating length (also applies to osmotic work)
l_T - 0 NA NA scaled heating length
ome - 4.01641 NA NA \omega, contribution of ash free dry mass of reserve to total ash free dry biomass
J_E_Am mol/d.cm^2 0.000628295 20 NA {J_EAm}, max surface-spec assimilation flux
y_E_X mol/mol 0.763636 NA NA yield of reserve on food
y_X_E mol/mol 1.30952 NA NA yield of food on reserve
p_Xm J/d.cm^2 431.953 20 NA {p_Xm}, max spec feeding power
J_X_Am mol/d.cm^2 0.000822767 20 NA {J_XAm}, max surface-spec feeding flux
y_P_X mol/mol 0.109375 NA NA yield of faeces on food
y_X_P mol/mol 9.14286 NA NA yield of food on faeces
y_P_E mol/mol 0.143229 NA NA yield of faeces on reserve
eta_XA mol/J 2.38095e-06 NA NA mass-power couplers for food on assimilation
eta_PA mol/J 2.60417e-07 NA NA mass-power couplers for product on assimlation
eta_VG mol/J 1.60044e-06 NA NA mass-power couplers for structure on growth
J_E_M mol/d.cm^3 0.000675695 20 NA [J_EM], vol-spec somatic maint costs
J_E_T mol/d.cm^2 0 20 NA {J_ET}, surface-spec somatic maint costs
j_E_M mol/d.mol 0.100932 20 NA mass-spec somatic maint costs
j_E_J mol/d.mol 0.0022721 20 NA mass-spec maturity maint costs
kap_G - 0.800222 NA NA \kappa_G, growth efficiency
E_V J/cm^3 3347.28 NA NA [E_V], volume-specific energy of structure
K mol X/l 0.00012658 NA NA half-saturation coefficient
M_Hb mol 2.56727e-07 NA NA maturity level at birth
U_Hb cm^2.d 0.00040861 20 NA scaled maturity level at birth
V_Hb cm^2.d 0.027277 20 NA scaled maturity level at birth
u_Hb - 4.32738e-05 NA NA scaled maturity level at birth
v_Hb - 0.00288877 NA NA scaled maturity level at birth
M_Hp mol 7.05091e-05 NA NA maturity level at puberty
U_Hp cm^2.d 0.112223 20 NA scaled maturity level at puberty
V_Hp cm^2.d 7.49152 20 NA scaled maturity level at puberty
u_Hp - 0.011885 NA NA scaled maturity level at puberty
v_Hp - 0.79339 NA NA scaled maturity level at puberty
t_E d 31.1198 22.5 NA maximum reserve residence time
t_starve d 31.593 22.5 0 maximum survival time when starved
s_M - 1 NA 1 acceleration factor at f=1
r_B 1/d 0.0083217 22.5 1 von Bertalanffy growth rate
W_dWm g 1.14207 NA 1 wet weight at maximum growth
dWm g/d 0.0179562 22.5 1 maximum growth in wet weight
U_E0 cm^2.d 0.0956387 20 1 scaled initial reserve
E_0 J 33.0491 NA 1 initial reserve
M_E0 mol 6.00893e-05 NA 1 initial reserve
Wd_0 g 0.00143613 NA 1 initial dry weight
Ww_0 g 0.00897583 NA 1 initial wet weight
l_b - 0.127482 NA 1 scaled structural length at birth
L_b cm 0.116763 NA 1 structural length at birth
M_Vb mol 1.06571e-05 NA 1 structural mass at birth
del_Ub - 0.712329 NA 1 fraction of reserve left at birth
Ww_b g 0.00798565 NA 1 wet weight at birth
Wd_b g 0.0012777 NA 1 dry weight at birth
E_Wb J 28.8704 NA 1 energy content at birth
a_b d 12.9594 22.5 1 age at birth
g_Hb - 0.40039 NA NA energy divestment ratio at birth
s_Hbp - 0.00364105 NA NA maturity ratio
s_HLbp - 0.32301 NA 1 maturity density ratio at f=1
l_p - 0.568562 NA 1 scaled structural length at puberty
L_p cm 0.520757 NA 1 structural length at puberty
M_Vp mol 0.000945428 NA 1 structural mass at puberty
M_Ep mol 0.00379723 NA 1 reserve mass at puberty
Ww_p g 0.708435 NA 1 wet weight at puberty
Wd_p g 0.11335 NA 1 dry weight at puberty
E_Wp J 2561.19 NA 1 energy content at puberty
a_p d 97.5887 22.5 1 age at puberty
g_Hp - 1.23956 NA NA energy divestment ratio at puberty
s_s - 0.000259589 NA 1 supply stress
l_i - 1 NA 1 ultimate scaled structural length
L_i cm 0.91592 NA 1 ultimate structural length
M_Vi mol 0.00514393 NA 1 ultimate structural mass
M_Ei mol 0.0206601 NA 1 ultimate reserve mass
Ww_i g 3.85448 NA 1 ultimate wet weight
Wd_i g 0.616717 NA 1 ultimate dry weight
E_Wi J 13935 NA 1 ultimate energy content
xi_WE kJ/ g 22.5955 NA 1 whole-body energy density of dry biomass (no reprod buffer)
del_Wb - 0.00207178 NA 1 birth weight as fraction of maximum weight
del_Wp - 0.183795 NA 1 puberty weight as fraction of maximum weight
del_V - 0.199346 NA 1 fraction of max weight that is structure
h_W 1/d 0.00034304 22.5 1 Weibull ageing rate
h_G 1/d 3.21339e-06 22.5 1 Gompertz aging rate
S_b - 1 NA 1 survival probability at birth
S_p - 0.999991 NA 1 survival probability at puberty
a_m d 2799.44 22.5 1 life span
a_99 d 549.965 22.5 1 age at length 0.99 * L_i
R_i 1/d 0.0772109 22.5 1 ultimate reproduction rate
N_i # 200.492 NA 1 life time reproductive output
M_E0_min_G mol 2.13956e-05 NA NA egg mass whereby growth ceases at birth
M_E0_min_R mol 1.77496e-05 NA NA egg mass whereby maturation ceases at birth
eb_min_G - 0.118014 NA NA scaled reserve density whereby growth ceases at birth
eb_min_R - 0.00610663 NA NA scaled reserve density whereby maturation ceases at birth
ep_min - 0.261394 NA NA scaled reserve density whereby maturation and growth cease at puberty
sM_min - 1 NA NA acceleration factor whereby maturation ceases at puberty
p_Xb J/d 7.41764 22.5 1 food intake at birth
J_Xb mol/d 1.41288e-05 22.5 1 food intake at birth
F_mb l/d 0.11162 22.5 1 max searching rate at birth
p_Xp J/d 147.545 22.5 1 food intake at puberty
J_Xp mol/d 0.000281039 22.5 1 food intake at puberty
F_mp l/d 2.22025 22.5 1 max searching rate at puberty
p_Xi J/d 456.425 22.5 1 ultimate food intake
J_Xi mol/d 0.000869381 22.5 1 ultimate food intake
F_mi l/d 6.86826 22.5 1 max ultimate searching rate
p_Ab J/d 5.93421 22.5 1 assimilation at birth
p_Cb J/d 1.91161 22.5 1 mobilisation at birth
p_Sb J/d 0.745178 22.5 1 somatic maintenance at birth
p_Jb J/d 0.000354799 22.5 1 maturity maintenance at birth
p_Gb J/d 1.1378 22.5 1 growth at birth
p_Rb J/d 0.0282811 22.5 1 maturation at birth
p_Db J/d 0.773813 22.5 1 dissipation at birth
p_Ap J/d 118.036 22.5 1 assimilation at puberty
p_Cp J/d 78.4719 22.5 1 mobilisation at puberty
p_Sp J/d 66.1056 22.5 1 somatic maintenance at puberty
p_Jp J/d 0.0974523 22.5 1 maturity maintenance at puberty
p_Gp J/d 11.1908 22.5 1 growth at puberty
p_Rp J/d 1.07806 22.5 1 reproduction at puberty
p_Dp J/d 67.2811 22.5 1 dissipation at puberty
p_Ai J/d 365.14 22.5 1 ultimate assimilation
p_Ci J/d 365.14 22.5 1 ultimate mobilisation
p_Si J/d 359.67 22.5 1 ultimate somatic maintenance
p_Ji J/d 0.0984979 22.5 1 ultimate maturity maintenance
p_Gi J/d 0 22.5 1 ultimate growth
p_Ri J/d 5.3713 22.5 1 ultimate reproduction
p_Di J/d 362.589 22.5 1 ultimate dissipation
J_Cb mol/d 3.19024e-06 22.5 1 CO2 flux at birth
J_Cp mol/d 0.000148421 22.5 1 CO2 flux at puberty
J_Ci mol/d 0.000711929 22.5 1 ultimate CO2 flux
J_Hb mol/d 3.0063e-06 22.5 1 water flux at birth
J_Hp mol/d 0.000139863 22.5 1 water flux at puberty
J_Hi mol/d 0.000670881 22.5 1 ultimate water flux
J_Ob mol/d 3.88002e-06 22.5 1 O2 flux at birth
J_Op mol/d 0.000180512 22.5 1 O2 flux at puberty
J_Oi mol/d 0.00086586 22.5 1 ultimate O2 flux
J_Nb mol/d 2.58668e-07 22.5 1 N-waste flux at birth
J_Np mol/d 1.20341e-05 22.5 1 N-waste flux at puberty
J_Ni mol/d 5.7724e-05 22.5 1 ultimate N-waste flux
RQ_b mol C/mol O 0.822222 NA 1 resp quotient at birth
UQ_b mol N/mol O 0.0666667 NA 1 urine quotient at birth
WQ_b mol H/mol O 0.733333 NA 1 water quotient at birth
SDA_b mol O/mol X 0.142862 NA 1 specific dynamic action at birth
VO_b L/h.g -0.00127017 22.5 1 dioxygen use per gram max dry weight, <J_OD> at birth
p_Tb J/d 1.7429 22.5 1 total heat at birth
RQ_p mol C/mol O 0.822222 NA 1 resp quotient at puberty
UQ_p mol N/mol O 0.0666667 NA 1 urine quotient at puberty
WQ_p mol H/mol O 0.733333 NA 1 water quotient at puberty
SDA_p mol O/mol X 0.142862 NA 1 specific dynamic action at puberty
VO_p L/h.g -0.00124489 22.5 1 dioxygen use per gram max dry weight, <J_OD> at puberty
p_Tp J/d 84.2713 22.5 1 total heat at puberty
RQ_i mol C/mol O 0.822222 NA 1 ultimate resp quotient
UQ_i mol N/mol O 0.0666667 NA 1 ultimate urine quotient
WQ_i mol H/mol O 0.733333 NA 1 ultimate water quotient
SDA_i mol O/mol X 0.142862 NA 1 ultimate specific dynamic action
VO_i L/h.g -0.00123306 22.5 1 ultimate dioxygen use per gram max dry weight, <J_OD>
p_Ti J/d 408.231 22.5 1 ultimate total heat