Vipera berus: Population traits

at T = 18 °C with background hazards for embryo, juv, adult: 0, 0, 0 1/d

Ln survival probability as function of age, with (dashed) and without (solid) thinning at max (red) and min (blue) scaled functional response for females.
Ln survivor function of the stable age distribution as function of age, with (dashed) and without (solid) thinning at max (red) and min (blue) scaled functional response for females.
model std f_minf f_max date: 2022/12/20
symbol units thin falsethin truethin falsethin truethin falsethin truedescription
f - 0.638022 0.652847 0.826423 0.826423 1 1 scaled functional response
r 1/d 0 0 0.00122851 0.00055539 0.00181833 0.000807567 spec pop growth rate
t2 d NaN NaN 564.216 1248.04 381.201 858.316 population doubling time
a_b d 219.707 218.917 211.25 211.25 205.637 205.637 age at birth
t_p d 4568.98 3365.45 1445.57 1445.57 1051.04 1051.04 time since birth at puberty
S_b - 0.999981 0.999981 0.999983 0.999983 0.999984 0.999984 survival probability at birth
S_p - 0.999868 0.659747 0.99989 0.586147 0.999903 0.534984 survival probability at puberty
theta_0b - 0.0265893 0.0930242 0.229123 0.298396 0.312135 0.379126 frac of ind that is embryo
theta_bp - 0.0266042 0.0750274 0.176814 0.20255 0.214819 0.234845 fraction of individuals that is juvenile
theta_pi - 0.946806 0.831948 0.594062 0.499054 0.473046 0.386029 frac of ind that is adult
L_bi cm 2.44462 2.36553 2.15906 2.16205 2.1541 2.15116 mean structural length of post-natals
L2_bi cm^2 6.06343 5.7777 4.99655 5.11726 5.06462 5.20344 mean squared structural length of post-natals
L3_bi cm^3 15.1922 14.4276 12.2809 13.0838 12.9081 14.0291 mean cubed structural length of post-natals
Ww_bi g 33.2874 32.0115 31.2279 33.2694 37.0055 40.2191 mean wet weight of post-natals
L_pi cm 2.47455 2.45488 2.37586 2.46394 2.46559 2.57631 mean structural length of adults
L2_pi cm^2 6.18017 6.1277 5.87092 6.37294 6.37777 7.07538 mean squared structural length of adults
L3_pi cm^3 15.5446 15.4919 15.0492 17.2154 17.2978 20.6543 mean cubed structural length of adults
Ww_pi g 34.0595 34.3729 38.2671 43.7754 49.5902 59.2126 mean wet weight of adults
R 1/d 0 0 0.000325802 0.00137741 0.00247856 0.00407932 mean reproduction rate of adults
J_X g/d 0.216328 0.210924 0.230904 0.236483 0.283208 0.290971 mean ingestion rate of wet food by post-natals
Y_PX mol/mol 0.109375 0.109375 0.109375 0.109375 0.109375 0.109375 yield of faeces on food
Y_VX mol/mol 0 0 0.0653399 0.0307277 0.0828759 0.0389365 yield of living structure on food
Y_VX_d mol/mol 0.0101336 0.0187304 0.000476505 0.0342955 0.000211376 0.0431359 yield of dead structure on food
Y_EX mol/mol 0 0 0.100807 0.0474068 0.154716 0.0726884 yield of living reserve on food
Y_EX_d mol/mol 0.01207 0.0228279 0.000735153 0.0529112 0.000394605 0.0805279 yield of dead reserve on food
Y_CX mol/mol 0.705592 0.689867 0.587654 0.589293 0.530097 0.532461 yield of CO2 on food
Y_HX mol/mol 0.75022 0.733499 0.624822 0.626564 0.563624 0.566138 yield of H2O on food
Y_OX mol/mol -0.89556 -0.8756 -0.745869 -0.747949 -0.672815 -0.675816 yield of O2 on food
Y_NX mol/mol 0.162829 0.1592 0.135613 0.135991 0.12233 0.122876 yield of N-waste on food
mu_TX J/mol 452849 442810 377126 378177 339676 341199 yield of heat on food

Remarks

  • Reprod-code O applies. Sex ratio is assumed to be 1:1. Parameters of male and female are the same.
  • Buffer handling rule: release each egg as soon as reproduction buffer allows
  • Weights of adults do not include reproduction buffer