Predictions & Data for this entry

Model: stf climate: MB migrate: phylum:
COMPLETE = 2.5 ecozone: MC food: biCi, biCvf class:
MRE = 0.092 habitat: 0iMcp gender: D order:
SMSE = 0.085 embryo: Mv reprod: O family:

Zero-variate data
ab408.8 380.1 (0.07016)dage at birthWass1973
tp4745 4107 (0.1345)dtime since birth at pubertyfishbase
am1.241e+04 1.241e+04 (5.754e-06)dlife spanfishbase
Lb 64 64.39 (0.006037)cmtotal length at birthWass1973
Lp154.9 186.2 (0.2019)cmfork length at puberty for femalesWiki
Lpm151.6 170.1 (0.1222)cmfork length at puberty for malesWiki
Li250 224.2 (0.1031)cmultimate total length for femalesfishbase
Lim180 204.2 (0.1347)cmultimate total length for malesfishbase
Wwi1.179e+05 1.161e+05 (0.01523)gultimate wet weightfishbase
Ri0.01507 0.01406 (0.06696)#/dmaximum reprod rateWass1973
Uni-variate data
DatasetFigure(RE)Independent variableDependent variableReference
tLe see Fig. 1 (0.195)agetotal lengthWass1973
tL_f see Fig. 2 (0.07142)time since birthtotal lengthRomiGrub2006
tL_m see Fig. 2 (0.07224)time since birthtotal lengthRomiGrub2006
Pseudo-data at Tref
DataGeneralised animalCarcharhinus plumbeusUnitDescription
v 0.02 0.03766cm/denergy conductance
kap 0.8 0.3568-allocation fraction to soma
kap_R 0.95 0.95-reproduction efficiency
p_M 18 5.415J/^3vol-spec som maint
k_J 0.002 0.00025821/dmaturity maint rate coefficient
kap_G 0.8 0.7977-growth efficiency
k 0.3 0.2501-maintenance ratio


  • egg development is initially fuelled by yolk only, later also absorb nutrients from fluids in the oviducts (ref: Wass1973)
  • Preferred temp 27 C,usually 20 - 65 m deep (ref: fishbase)


  • Males are assumed to differ from females by {p_Am} only
  • In view of low somatic maintenance, pseudodata k_J = 0.002 1/d is replaced by pseudodata k = 0.3


  • [Wiki]
  • [fishbase]
  • [Kooy2010] S.A.L.M. Kooijman. Dynamic Energy Budget theory for metabolic organisation. Cambridge Univ. Press, Cambridge, 2010.
  • [RomiGrub2006] J. G. Romine, R. D. Grubbs, and J. A. Musick. Age and growth of the sandbar shark, Carcharhinus plumbeus, in Hawaiian waters through vertebral analysis. Environ Biol Fishs, 77:229--239, 2006.
  • [Wass1973] R. C. Wass. Size, growth, and reproduction of the sandbar shark, Carcharhinus milberti, in Hawaii. Pacific Science, 27(4):305--318, 1973.

Bibtex file with references for this entry

Bas Kooijman, 2017/08/03

accepted: 2017/08/03

refer to this entry as: AmP Carcharhinus plumbeus version 2017/08/03