Predictions & Data for this entry

Model: hep climate: Cfb, Dfb, Dfc migrate:
COMPLETE = 2.5 ecozone: TH food: beH, beD, beS
MRE = 0.189 habitat: 0eFr, 0eFc, eiTg gender: D
SMSE = 0.277 embryo: Fs reprod: O

Zero-variate data
ab 25 21.79 (0.1282)dage at birthHump1984
tj 90 82.52 (0.08313)dtime since at birth at emergenceHump1984
am 2 2.039 (0.0193)dlife span as imagoguess
Lj 1 0.7536 (0.2464)cmmax body length nymphHump1981
Li1.5 1.506 (0.004157)cmbody length imagoMacaBenn2010
Wd00.0006 0.0006155 (0.0259)mginitial dry weight Wata1998, guess
Wdj 2 2.371 (0.1857)mgultimate dry weightHump1981
Ni2500 2010 (0.1961)#cumulative # of eggs riverflies, MacaBenn2010
Uni-variate data
DatasetFigure(RE)Independent variableDependent variableReference
tL_20 see Fig. 1 (0.158)timebody lengthHump1981
tL_14 see Fig. 1 (0.08538)timebody lengthHump1981
tL_9 see Fig. 1 (0.1829)timebody lengthHump1981
tW_20 see Fig. 2 (0.1036)timedry weightHump1981
tW_14 see Fig. 2 (0.3844)timedry weightHump1981
tW_9 see Fig. 2 (0.7093)timedry weightHump1981
Tab see Fig. 3 (0.3169)temperatureage at birthHump1984
Pseudo-data at Tref
DataGeneralised animalEcdyonurus disparUnitDescription
v 0.02 0.002135cm/denergy conductance
kap 0.8 0.4018-allocation fraction to soma
kap_R 0.95 0.95-reproduction efficiency
p_M 18 19.65J/^3vol-spec som maint
k_J 0.002 0.0021/dmaturity maint rate coefficient
kap_G 0.8 0.8083-growth efficiency


  • Imagos are extremely short lived and do not feed; nyphs allocate to reproduction buffer and have mature eggs (ref: Sold1979)
  • Estimates of the number of instars in ephemeropterans vary between 10 and 50; most are in the range 15-25 (ref: Brit1982)
  • Final emphemeropteran larval stage transforms into a flying sub-image, which moulds into an imago (ref: Brit1982)
  • eggs have a dormant period; length grows by a fixed fraction over moulting (so Dyar rule is applicable); eggs can develop parthenogenetically (ref: Hump1981)


  • Puberty is assumed to coincide with end of acceleration


  • [Wiki]
  • [riverflies]
  • [Brit1982] J. E. Brittain. Biology of mayflies. Ann. Rev. Entomol., 27:119--147, 1982.
  • [Hump1981] U. H. Humpesch. Effect of temperature on larval growth of Ecdyonurus dispar (Ephemeroptera: Heptageniidae) from two English lakes. Freshwater Biol., 11:441--457, 1981.
  • [Hump1984] U. H. Humpesch. Egg development of non-diapausing exopterygote aquatic insects occurring in Europe. 1984.
  • [Kooy2010] S.A.L.M. Kooijman. Dynamic Energy Budget theory for metabolic organisation. Cambridge Univ. Press, Cambridge, 2010.
  • [MacaBenn2010] C. Macadam and C. Bennet. A pictorial guide to British Ephemeroptera. Field Studies Council, 2010.
  • [Sold1979] T. Soldan. The structure and development of the female internal reproductive system in six European species of Ephemeroptera. Acta entomol. bohemoslovaca, 76:353--365, 1979.
  • [Wata1998] N. C. W. Watanabe. Geographical variation in japan in egg development of the mayfly, Ephoron shigae (Ephemeroptera: Polymitarcyidae). Freshwater Biol., 40:245--254, 1981.

Bibtex file with references for this entry

Bas Kooijman, 2014/10/29 (last modified by Bas Kooijman 2016/02/11)

accepted: 2016/02/19

refer to this entry as: AmP Ecdyonurus dispar version 2016/02/19