Predictions & Data for this entry

Model: abj climate: A, B, C, D migrate:
COMPLETE = 1.7 ecozone: TH, TN, TP, TA food: biHa, biCi, biB
MRE = 0.124 habitat: 0iT gender: D
SMSE = 0.153 embryo: Tns reprod: Apf

Zero-variate data
ab 7 6.469 (0.0758)dtime since t_0 at birthGrev1980
tp 24 30.24 (0.2599)dtime since birth at pubertyGrev1980
am900 900.4 (0.0004712)dlife spanEarth
Lb0.02 0.02164 (0.08193)cmtotal length at birthGrev1980
Lp0.0435 0.03644 (0.1622)cmtotal length at pubertyGrev1980
Li0.1 0.1045 (0.04489)cmultimate total lengthMorgKing1976
Wwb4.25e-07 4.773e-07 (0.1232)gwet weight at birthestimate
Wwp4.37e-06 2.28e-06 (0.4782)gwet weight at pubertyestimate
Wwi5.31e-05 5.375e-05 (0.01218)gultimate wet weightestimate
Ri0.1375 0.1377 (0.001445)#/dmaximum reprod ratewild guess
Pseudo-data at Tref
DataGeneralised animalMilnesium tardigradumUnitDescription
v 0.02 0.005628cm/denergy conductance
kap 0.8 0.9305-allocation fraction to soma
kap_R 0.95 0.95-reproduction efficiency
p_M 18 40.13J/^3vol-spec som maint
k_J 0.002 0.0021/dmaturity maint rate coefficient
kap_G 0.8 0.8109-growth efficiency


  • The cuticle is moulted about four times during the life cycle. (ref: bumblebee)
  • Some species lay their eggs inside the moulted cuticle, into which males shed sperm. (ref: bumblebee)
  • It appears that egg laying and defecation can only happen during moulting. (ref: bumblebee)


  • [Wiki]
  • [bumblebee]
  • [Earth]
  • [Baum1964] H. Baumann. Uber den Lebenslauf und die Lebensweise von Milnesium tardigradum Doyere (Tardigrada). Veröffentlichungen aus dem Überseemuseum Bremen, 3:161--171, 1964.
  • [Grev1980] H. Greven. Die Baertierchen, volume 537 of Die Neue Brehm-Bucherei. A. Ziemsen Verlag, Wittenberg Lutherstadt, 1980.
  • [Kooy2010] S.A.L.M. Kooijman. Dynamic Energy Budget theory for metabolic organisation. Cambridge Univ. Press, Cambridge, 2010.
  • [MorgKing1976] C. I. Morgan and P. E. King. British Tardigrades, volume 9 of Synopsis of the British Fauna. Academic Press, London, 1976.

Bibtex file with references for this entry

Bas Kooijman, 2012/12/24 (last modified by Bas Kooijman 2015/03/12)

accepted: 2016/03/02

refer to this entry as: AmP Milnesium tardigradum version 2016/03/02