Predictions & Data for this entry

Model: abj climate: Cfa, Cfb, Dfa migrate:
COMPLETE = 2.6 ecozone: THp food: biCi
MRE = 0.098 habitat: 0iFr, 0iFc gender: D
SMSE = 0.105 embryo: Fg reprod: O

Zero-variate data
ah_8 22 27.07 (0.2303)dage at hath at 8CIngrIbbo20aa
ah_1314.7 12.75 (0.1326)dage at hath at 13CIngrIbbo20aa
ah_15 8 9.504 (0.188)dage at hath at 15CIngrIbbo20aa
ap365 350.3 (0.04014)dage at pubertyLaak2008
am5110 5243 (0.026)dlife span IngrIbbo20aa
Lh1.7 1.086 (0.3612)cmtotal length at hatchIngrIbbo20aa
Lp 23 24.27 (0.05522)cmtotal length at pubertyLaak2008
Li 60 63.21 (0.05342)cmultimate total length fishbase
R4599.29 90.19 (0.09163)#/dreprod rate at 45 cm, 1160 g, 9 yr Laak2008
GSI0.1667 0.1672 (0.002912)-Gonado som index Laak2008
Uni-variate data
DatasetFigure(RE)Independent variableDependent variableReference
tL_Test see Fig. 1 (0.104)time since birthtotal(?) length Hell1969
tL_Storsjo see Fig. 1 (0.02832)time since birthtotal(?) length Hell1969
tL_Indals see Fig. 1 (0.02459)time since birthtotal(?) length Hell1969
tL_Llyn see Fig. 1 (0.03419)time since birthtotal(?) length Hell1969
tW see Fig. 2 (0.1247)time since birthwet weight Hell1969
LW see Fig. 3 (0.09194)total(?) lengthwet weight Hell1969
LW1 see Fig. 4 (0.02181)fork lengthwet weight Laak2008
Tah see Fig. 5 (0.1501)temperatureage at hatch JungWink1984
Pseudo-data at Tref
DataGeneralised animalThymallus thymallusUnitDescription
v 0.02 0.1542cm/denergy conductance
kap 0.8 0.3881-allocation fraction to soma
kap_R 0.95 0.95-reproduction efficiency
p_M 18 85.29J/^3vol-spec som maint
k_J 0.002 0.0021/dmaturity maint rate coefficient
kap_G 0.8 0.804-growth efficiency


  • 30.5 cm total length corresponds with 27.0 cm fork length (ref: Clar1992)
  • eggs are 3-4 mm, but 2 mm is also possible (ref: Laak2008)
  • ab-ah is 2 till 5 d (ref: Laak2008)
  • Clar1992: Lp = 28 cm (ref: Clar1992)


  • [Wiki]
  • [fishbase]
  • [Clar1992] R. A. Clark. Age and size at maturity of Arctic grayling in selected waters of the Tanana drainage., 1992.
  • [Hell1969] J. M. Hellawell. Age determination and growth of the grayling Thymallus thymallus (L.) of the River Lugg, Herefordshire. J Fish Biol, 1:373--382, 1969.
  • [IngrIbbo20aa] A. Ingram, A. Ibbotson, and M. Gallagher. The ecology and managements of the European grayling Thymallus thymallus (Linnaeus). Technical report, Institute of freshwater ecology, 20aa.
  • [JungWink1984] M. Jungwirth and H. Winkler. The temperature dependence of embryonic development og grayling (Thymallus, thymallus), Danube salmon (Hucho hucho), Arctic char (Salvelinus alpinus) and brown trout (Salmo trutta fario). Aquaculture, 38:315--327, 1984.
  • [Kooy2010] S.A.L.M. Kooijman. Dynamic Energy Budget theory for metabolic organisation. Cambridge Univ. Press, Cambridge, 2010.
  • [Laak2008] G. A. J. de Laak. Vlagzalm Thymallus thymallus (Linnaeus, 1758). Technical report, Sportvisserij Nederland, Postbus 162, 3720 AD Bilthoven, 2008.

Bibtex file with references for this entry

Bas Kooijman, 2012/04/30 (last modified by Bas Kooijman 2016/10/18)

accepted: 2016/10/18

refer to this entry as: AmP Thymallus thymallus version 2016/10/18