Predictions & Data for this entry

Model: abj climate: MC migrate: phylum:
COMPLETE = 2.8 ecozone: MAN food: bjPp, bjD class:
MRE = 0.177 habitat: 0jMc, jiMb gender: D order:
SMSE = 0.273 embryo: Mp reprod: O family:

Zero-variate data
ab1.667 1.84 (0.1043)dage at emergenceGoul1969
tj58.33 9.785 (0.8322)dtime since birth at metamGoul1969
tp547.5 472.4 (0.1371)dtime since birth at pubertySuer1984
am3650 3650 (4.043e-05)dlife spanSuer1984
Wwb8.2e-06 7.706e-06 (0.0602)gwet weight at birthguess
Wwj0.0002 0.0002093 (0.0467)gwet weight at settleSuer1984
Wwp 56 58.86 (0.05099)gwet weight at pubertySuer1984
Wwi160 151.4 (0.0539)gultimate wet weightSuer1984
Ri1.096e+05 1.004e+05 (0.08353)#/dreprod rateguess
Uni-variate data
DatasetFigure(RE)Independent variableDependent variableReference
V0V1_A see Fig. 1 (0.2939)initial volumefinal volumeSuer1984
V0V1_S see Fig. 1 (0.2795)initial volumefinal volumeSuer1984
Pseudo-data at Tref
DataGeneralised animalUrechis caupoUnitDescription
v 0.02 0.05545cm/denergy conductance
kap 0.8 0.6-allocation fraction to soma
kap_R 0.95 0.95-reproduction efficiency
p_M 18 2J/^3vol-spec som maint
k_J 0.002 0.00054941/dmaturity maint rate coefficient
kap_G 0.8 0.7887-growth efficiency
k 0.3 0.2915-maintenance ratio


  • Suspension feeder, contrary to e.g. Echiurus, which is a deposit feeder (ref: Suer1984)


  • Adjusted d_V and d_E of 0.04 g/cm^3 is used
  • In absence of reprod data, kap = 0.6 is fixed for this high-fertility species
  • Spec som maint is fixed at 2 J/^3, since par-estimation results in tiny values that are hard to believe
  • Under-estimation of t_j might be linked to veliating assimilation by trochophora larvae
  • In view of low somatic maintenance, pseudodata k_J = 0.002 1/d is replaced by pseudodata k = 0.3


  • [Wiki]
  • [Goul1969] M. C. Gould. A comparison of RNA and protein synthesis in fertilized and unfertilized eggs of Urechis caupo. Developmental Biology, 19:482--497, 1969.
  • [Kooy2010] S.A.L.M. Kooijman. Dynamic Energy Budget theory for metabolic organisation. Cambridge Univ. Press, Cambridge, 2010.
  • [Suer1984] A. L. Suer. Growth and spawning of Urechis caupo (Echiura) in Bodega Harbor, California. Marine Biology, 78:275--284, 1984.

Bibtex file with references for this entry

Bas Kooijman, 2017/09/20

accepted: 2017/09/20

refer to this entry as: AmP Urechis caupo version 2017/09/20