Predictions & Data for this entry

Model: abj climate: MC migrate:
COMPLETE = 2.1 ecozone: MAE food: biCi
MRE = 0.112 habitat: 0iMcd gender: D
SMSE = 0.101 embryo: Mv reprod: O

Zero-variate data
ab 90 85.08 (0.05463)dage at birthVete1999
ap265 423.8 (0.5994)dage at puberty Brod2001, Vete1999
am111460 1615 (0.1061)dlife spanBrod2001
am42920 2647 (0.09335)dlife spanBrod2001
Lb5.1 4.983 (0.02297)cmtotal length at birthVete1999
Lp21.5 18.67 (0.1314)cmtotal length at puberty for femaleVete1999
Lpm 19 16.02 (0.1569)cmtotal length at puberty for maleVete1999
Li 50 51.75 (0.03496)cmultimate total length for femaleVeer2009
Lim 42 43.5 (0.03576)cmultimate total length for maleVeer2009
rB110.001 0.001102 (0.1021)1/dvon Bertalanffy growth rateBrod2001
rB40.0006 0.0006723 (0.1205)1/dvon Bertalanffy growth rateBrod2001
Wwb0.46 0.4829 (0.04969)gwet weight at birthVete1999
Wwp27.4 25.42 (0.07234)gwet weight at puberty for femaleVete1999
Wwpm13.7 13.8 (0.007552)gwet weight at puberty for maleVete1999
Wwi510 540.8 (0.06046)gultimate wet weight for femalefishbase
Wwim315 321.3 (0.02002)gultimate wet weight for maleVeer2009
Ww33153 140.3 (0.08327)gwet weight at 33 cmVete1999
R200.06575 0.07977 (0.2131)#/dreprod rate at 20 cmWitt2009
R330.3425 0.2843 (0.17)#/dreprod rate at 33 cmVete1999
Pseudo-data at Tref
DataGeneralised animalZoarces viviparusUnitDescription
v 0.02 0.03059cm/denergy conductance
kap 0.8 0.667-allocation fraction to soma
kap_R 0.95 0.95-reproduction efficiency
p_M 18 169.1J/^3vol-spec som maint
k_J 0.002 0.0021/dmaturity maint rate coefficient
kap_G 0.8 0.8051-growth efficiency


  • Longest known pregnancy among fish (ref: Wiki)


  • Males are assumed to differ from females by {p_Am} only
  • abj has been applied since related species show acceleration; foetal development was ignored because of absence of its combination with abj. The effect is probably very small.


  • [Wiki]
  • [fishbase] url{}.
  • [Witt2009] Hans Witte (NIOZ), pers. comm.
  • [Veer2009] Hanek van der Veer (NIOZ), pers. comm.
  • [Brod2001] E. Brodte. Wachstum und Fruchtbarkeit der Aalmutterarten Zoarces viviparus (L.) und Pachycara brachycephalum (Pappenheim) aus unterschiedlichen klimatischen Regionen. PhD thesis, University of Bremen, 2001.
  • [Kooy2010] S.A.L.M. Kooijman. Dynamic Energy Budget theory for metabolic organisation. Cambridge Univ. Press, Cambridge, 2010.
  • [Vete1999] M. Vetemaa. Reproduction biology of the viviparous blenny (Zoarces viviparus L.) in Kattegat. Fiskeriverket rapport, 1999:81--96, 1999.

Bibtex file with references for this entry

Vania Freitas, 2009/07/09 (last modified by Bas Kooijman 2016/10/15)

accepted: 2016/10/15

refer to this entry as: AmP Zoarces viviparus version 2016/10/15